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The elements off the diagonal represent the additive genetic relationship between individual i and other individual j (1 ≤ i, j ≤ 2N-1).
These elements are calculated
from the multivariate disequilibrium coefficients C;
through equations Cij = α1i, α2j + α22i, α21j and α11j.
where α11 (that is, the diagonal element corresponding to the origin) equals 1, α21 equals the inbreeding coefficient of individual ec5d62056f elitzym
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